Datasets

Glycine datasets

Glycine max

Genomes

58-161.gnm1.BW8J
Genome assembly for Glycine max accession 58-161 (SoyL04)
Amsoy.gnm1.TYYG
Genome assembly for Glycine max accession Amsoy (SoyC05)
DongNongNo_50.gnm1.XRTN
Genome assembly for Glycine max accession DongNongNo_50 (SoyC12)
FengDiHuang.gnm1.YYLM
Genome assembly for Glycine max accession FengDiHuang (SoyL07)
FiskebyIII.gnm1.F177
Glycine max genotype Fiskeby III genome assembly 1; JGI name Fiskeby v1.1
HanDouNo_5.gnm1.11MF
Genome assembly for Glycine max accession HanDouNo_5 (SoyC09)
Hefeng25_IGA1002.gnm1.L69T
Genome assembly files for cultivar Hefeng 25 (Hefeng25_IGA1002 in publication; WHFS_GmHF25_1.0 in the GenBank assembly record)
HeiHeNo_43.gnm1.FV4H
Genome assembly for Glycine max accession HeiHeNo_43 (SoyC13)
Huaxia3_IGA1007.gnm1.RGGN
Genome assembly files for cultivar Huaxia3 (Huaxia3_IGA1007 in publication; WHFS_GmHX3_1.0 in the GenBank assembly record)
Hwangkeum.gnm1.4S83
Glycine max genotype Hwangkeum genome assembly v1.0
JD17.gnm1.TW7L
Glycine max accession Jidou 17 (JD17), genome assembly 1
JiDouNo_17.gnm1.MRYM
Genome assembly for Glycine max accession JiDouNo_17 (SoyC11)
JinDouNo_23.gnm1.GBYD
Genome assembly for Glycine max accession JinDouNo_23 (SoyC07)
Jinyuan_IGA1006.gnm1.LXM0
Genome assembly files for cultivar Jinyuan (Jinyuan_IGA100 in the publication; 6HFS_GmJY_1.0 in the GenBank assembly record)
JuXuanNo_23.gnm1.WC57
Genome assembly for Glycine max accession JuXuanNo_23 (SoyC03)
KeShanNo_1.gnm1.3TDD
Genome assembly for Glycine max accession KeShanNo_1 (SoyC14)
Lee.gnm1.BXNC
Glycine max accession Lee Genome assembly 1; JGI name Lee v1.1
Lee.gnm2.K7BV
Glycine max genotype Lee genome assembly v2.0
Lee.gnm3.VG1C
Glycine max accession Lee, genome assembly 3
PI_398296.gnm1.D07C
Genome assembly for Glycine max accession PI_398296 (SoyL05)
PI_548362.gnm1.V9S7
Genome assembly for Glycine max accession PI_548362 (SoyC10)
QiHuangNo_34.gnm1.MB9G
Genome assembly for Glycine max accession QiHuangNo_34 (SoyC08)
ShiShengChangYe.gnm1.MC01
Genome assembly for Glycine max accession ShiShengChangYe (SoyL09)
TieFengNo_18.gnm1.BW65
Genome assembly for Glycine max accession TieFengNo_18 (SoyC02)
TieJiaSiLiHuang.gnm1.W9PV
Genome assembly for Glycine max accession TieJiaSiLiHuang (SoyL08)
TongShanTianEDan.gnm1.B4R6
Genome assembly for Glycine max accession TongShanTianEDan (SoyL03)
WanDouNo_28.gnm1.ZRP2
Genome assembly for Glycine max accession WanDouNo_28 (SoyC04)
Wenfeng7_IGA1001.gnm1.L0QH
Genome assembly files for cultivar Wenfeng 7 (Wenfeng7_IGA1001 in publication; WHFS_GmWF7_1.0 in the GenBank assembly record); Chu et al. (2021)
Wm82.gnm1.FCtY
Glycine max accession Williams genome assembly v1.0; JGI name Glycine max v1.1
Wm82.gnm2.DTC4
Glycine max accession Williams 82 genome assembly v2.0; JGI name Wm82.a2.v1
Wm82.gnm4.4PTR
Glycine max accession Williams 82 genome assembly v4.0; JGI name Wm82.a4.v1
Wm82.gnm5.NRKG
Glycine max accession Williams 82 (Wm82), genome assembly 5
Wm82.gnm6.S97D
Glycine max accession Williams 82 (ISU01) genome assembly v6; renamed from Wm82 ISU-01 v2.1; JGI name Wm82.a6.v1
Wm82_IGA1008.gnm1.5CQQ
Genome assembly files for cultivar Williams 82 (Wm82_IGA1008 in publication; WHFS_GmW82_1.0 in the GenBank assembly record)
Wm82_NJAU.gnm1.N4GV
Glycine max accession Williams 82 from Nanjing Agricultural University (Wm82-NJAU), genome assebly v1
XuDouNo_1.gnm1.0GWH
Genome assembly for Glycine max accession XuDouNo_1 (SoyC01)
YuDouNo_22.gnm1.T2VD
Genome assembly for Glycine max accession YuDouNo_22 (SoyC06)
Zh13.gnm1.N6C8
Genome assembly files for cultivar Zhonghuang 13, Shen et al. (2018)
Zh13.gnm2.LV9P
Genome assembly version 2 files for cultivar Zhonghuang 13, Shen et al. (2019)
Zh13_IGA1005.gnm1.FRXQ
Genome assembly files for cultivar Zhonghuang 13 (Zh13_IGA1005 in publication; WHFS_GmZH13_1.0 in the GenBank assembly record)
Zh35_IGA1004.gnm1.DBYJ
Genome assembly files for cultivar Zhonghuang 35 (Zh35_IGA1004 in publication; WHFS_GmZH35_1.0 in the GenBank assembly record)
ZhangChunManCangJin.gnm1.FC4Q
Genome assembly for Glycine max accession ZhangChunManCangJin (SoyL06)
Zhutwinning2.gnm1.GR6N
Genome assembly for Glycine max accession Zhutwinning2 (SoyL01)
ZiHuaNo_4.gnm1.HN60
Genome assembly for Glycine max accession ZiHuaNo_4 (SoyL02)

Annotations

58-161.gnm1.ann1.HJ1K
Gene annotations for Glycine max accession 58-161 (SoyL04)
Amsoy.gnm1.ann1.6S5P
Gene annotations for Glycine max accession Amsoy (SoyC05)
DongNongNo_50.gnm1.ann1.QSDB
Gene annotations for Glycine max accession DongNongNo_50 (SoyC12)
FengDiHuang.gnm1.ann1.P6HL
Gene annotations for Glycine max accession FengDiHuang (SoyL07)
FiskebyIII.gnm1.ann1.SS25
Genome annotation files for Glycine max genotype Fiskeby III, Stupar et al. (2020); JGI name Fiskeby v1.1
HanDouNo_5.gnm1.ann1.ZS7M
Gene annotations for Glycine max accession HanDouNo_5 (SoyC09)
Hefeng25_IGA1002.gnm1.ann1.320V
Genome annotation files for cultivar Hefeng 25, Chu et al (2021)
HeiHeNo_43.gnm1.ann1.PDXG
Gene annotations for Glycine max accession HeiHeNo_43 (SoyC13)
Huaxia3_IGA1007.gnm1.ann1.LKC7
Genome annotation files for cultivar Huaxia3
Hwangkeum.gnm1.ann1.1G4F
Genome annotation files for Glycine max genotype Hwangkeum, Kim et al. (2021)
JD17.gnm1.ann1.CLFP
Annotation 1 for Glycine max accession Jidou 17 (JD17), genome assembly 1
JiDouNo_17.gnm1.ann1.X5PX
Gene annotations for Glycine max accession JiDouNo_17 (SoyC11)
JinDouNo_23.gnm1.ann1.SGJW
Gene annotations for Glycine max accession JinDouNo_23 (SoyC07)
Jinyuan_IGA1006.gnm1.ann1.2NNX
Genome annotation files for cultivar Jinyuan
JuXuanNo_23.gnm1.ann1.H8PW
Gene annotations for Glycine max accession JuXuanNo_23 (SoyC03)
KeShanNo_1.gnm1.ann1.2YX4
Gene annotations for Glycine max accession KeShanNo_1 (SoyC14)
Lee.gnm1.ann1.6NZV
Glycine max accession Lee genome annotation files; JGI name Lee v1.1
Lee.gnm2.ann1.1FNT
genome annotation of Glycine max Lee, genome assembly 2
Lee.gnm3.ann1.ZYY3
Annotation 1 for Glycine max accession Lee, genome assembly 3
PI_398296.gnm1.ann1.B0XR
Gene annotations for Glycine max accession PI_398296 (SoyL05)
PI_548362.gnm1.ann1.LL84
Gene annotations for Glycine max accession PI_548362 (SoyC10)
QiHuangNo_34.gnm1.ann1.WHRV
Gene annotations for Glycine max accession QiHuangNo_34 (SoyC08)
ShiShengChangYe.gnm1.ann1.VLGS
Gene annotations for Glycine max accession ShiShengChangYe (SoyL09)
TieFengNo_18.gnm1.ann1.7GR4
Gene annotations for Glycine max accession TieFengNo_18 (SoyC02)
TieJiaSiLiHuang.gnm1.ann1.W70Z
Gene annotations for Glycine max accession TieJiaSiLiHuang (SoyL08)
TongShanTianEDan.gnm1.ann1.56XW
Gene annotations for Glycine max accession TongShanTianEDan (SoyL03)
WanDouNo_28.gnm1.ann1.NLYP
Gene annotations for Glycine max accession WanDouNo_28 (SoyC04)
Wenfeng7_IGA1001.gnm1.ann1.ZK5W
Genome annotation files for cultivar Wenfeng 7
Wm82.gnm1.ann1.DvBy
Glycine max accession Williams genome annotation files; JGI name Glycine max v1.1
Wm82.gnm2.ann1.RVB6
Glycine max accession Williams genome annotation files; JGI name Wm82.a2.v1
Wm82.gnm4.ann1.T8TQ
Glycine max accession Williams genome annotation files; JGI name Wm82.a4.v1
Wm82.gnm5.ann1.J7HW
Annotation 1 for Glycine max accession Williams 82 (Wm82), genome assembly 5
Wm82.gnm6.ann1.PKSW
Annotation 1 Glycine max accession Williams 82 (ISU01) genome assembly v6; renamed from Wm82 ISU-01 v2.1; JGI name Wm82.a6.v1
Wm82_IGA1008.gnm1.ann1.FGN6
Genome annotation files for cultivar William 82
Wm82_NJAU.gnm1.ann1.KM71
Annotation 1 for Glycine max accession Williams 82 from Nanjing Agricultural University (Wm82-NJAU), genome assembly v1
XuDouNo_1.gnm1.ann1.G2T7
Gene annotations for Glycine max accession XuDouNo_1 (SoyC01)
YuDouNo_22.gnm1.ann1.HCQ1
Gene annotations for Glycine max accession YuDouNo_22 (SoyC06)
Zh13.gnm1.ann1.8VV3
Genome annotation files for cultivar Zhonghuang 13, Shen et al. (2018)
Zh13.gnm2.ann1.FJ3G
Genome annotation files for cultivar Zhonghuang 13, Shen et al. (2019)
Zh13_IGA1005.gnm1.ann1.87Z5
Genome annotation files for cultivar Zhonghuang 13
Zh35_IGA1004.gnm1.ann1.RGN6
Genome annotation files for cultivar Zhonghuang 35, Chu et al. (2021)
ZhangChunManCangJin.gnm1.ann1.7HPB
Gene annotations for Glycine max accession ZhangChunManCangJin (SoyL06)
Zhutwinning2.gnm1.ann1.ZTTQ
Gene annotations for Glycine max accession Zhutwinning2 (SoyL01)
ZiHuaNo_4.gnm1.ann1.FCFQ
Gene annotations for Glycine max accession ZiHuaNo_4 (SoyL02)

Expression

Wm82.gnm2.ann1.expr.Wm82.Libault_Farmer_2010
Soybean gene expression atlas from 14 conditions (Libault et al., 2010).
Wm82.gnm6.ann1.expr.Magellan.Pelaez-Vico_Sinha_2023
Combined phenotypic and transcriptomic analysis of soybean plants subjected to multifactorial stress combination (Pelaez-Vico et al.)
Wm82.gnm6.ann1.expr.Wm82.Chen_Lin_2021
Gene Expression Changes in the Development of the Soybean Seed-Heart Stage.
Wm82.gnm6.ann1.expr.Wm82.Chen_Lin_2021_2
Gene Expression Changes in the Development of the Soybean Seed-Globular Stage.
Wm82.gnm6.ann1.expr.Wm82.Chen_Lin_2021_3
Gene Expression Changes in the Development of the Soybean Seed-Cotyledon Stage.
Wm82.gnm6.ann1.expr.Wm82.Libault_Farmer_2010
Time-series transcriptomic data with Bradyrhizobium japonicum infection and mock inoculation
Wm82.gnm6.ann1.expr.Wm82.Sreedasyam_Plott_2023
Transcriptome sequencing as part of the JGI Flagship Plant Gene Atlas Inititative
Wm82.gnm6.ann1.expr.mixed.Kour_Boone_2014
Expression profiling by high throughput sequencing

Markers

FiskebyIII.gnm1.mrk.SoySNP50K
SoySNP50K is an Illumina Infinium II BeadChip that contains over 50,000 SNPs from soybean useful for genotyping a wide variety of cultivated and wild soybean accessions.
Wm82.gnm1.mrk.BARCSOYSSR
BARCSOYSSR contains 33,065 locus-specific SSR markers with di-, tri-, and tetranucleotide repeats of five or more.
Wm82.gnm1.mrk.Li_Guo_2016
59,845 single-nucleotide polymorphisms identified from re-sequencing of 279 accessions from Yangtze-Huai soybean breeding germplasm.
Wm82.gnm1.mrk.SoySNP50K
SoySNP50K is an Illumina Infinium II BeadChip that contains over 50,000 SNPs from soybean useful for genotyping a wide variety of cultivated and wild soybean accessions.
Wm82.gnm1.mrk.SoySNP6K
Illumina Infinium SoySNP6K BeadChip
Wm82.gnm1.mrk.SoySSR
This marker set contains SSRs developed and published by Cregan, et al. in 1999, with names starting with Satt or Sat_.
Wm82.gnm1.mrk.Zhao_Teng_2017
SLAF-seq markers associated with resistance to soybean cyst nematode (SCN) HG type 2.5.7, with respect to Wm82 gnm1 coordinates.
Wm82.gnm2.mrk.1536_USLP1
Universal Soy Linkage Panel (USLP 1.0) contains 1536 SNPs selected based on even distribution throughout each of the 20 consensus linkage groups and to have a broad range of allele frequencies in diverse germplasm.
Wm82.gnm2.mrk.BARCSOYSSR
BARCSOYSSR contains 33,065 locus-specific SSR markers with di-, tri-, and tetranucleotide repeats of five or more.
Wm82.gnm2.mrk.Fang_Ma_2017
GWAS of 84 agronomic traits, conducted on 809 soybean accessions, with genotyping by resequencing.
Wm82.gnm2.mrk.Han_Zhao_2015
SNP markers generated Specific Locus Amplified Fragment Sequencing (SLAF-seq) approach, and showing association with resistance against two soybean cyst nematode types.
Wm82.gnm2.mrk.Li_Zhao_2019
12,072 SNPs from Li, Zhao, et al. (2019), a seed protein and oil study.
Wm82.gnm2.mrk.Meng_He_2016
GWAS study of seed isoflavone content (SIFC), conducted on 366 landraces, genotyped with 116,769 RAD-Seq markers.
Wm82.gnm2.mrk.NJAU355K
NJAU 355K SoySNP array used to study soybean domestication in Wang, et al. (2016).
Wm82.gnm2.mrk.Sonah_ODonoughue_2015
GWAS study of eight traits in soybean, conducted on 139 accessions, genotyped using GBS-derived markers.
Wm82.gnm2.mrk.SoySNP50K
SoySNP50K is an Illumina Infinium II BeadChip that contains over 50,000 SNPs from soybean useful for genotyping a wide variety of cultivated and wild soybean accessions.
Wm82.gnm2.mrk.SoySNP6K
Illumina Infinium SoySNP6K BeadChip
Wm82.gnm2.mrk.SoySSR
This marker set contains SSRs developed and published by Cregan, et al. in 1999, with names starting with Satt or Sat_.
Wm82.gnm2.mrk.SoyaSNP180K
Axiom SoyaSNP180K array with 180,981 SNPs developed by Lee, et al. (2015).
Wm82.gnm2.mrk.Tran_Steketee_2019
Diverse soybean accessions (462) were screened for resistance to SCN and genotyped using SoySNP50K Infinium Chip and a set of 3 funtional KASP SNP markers, identifying both 88788-type and Peking-type resistance.
Wm82.gnm2.mrk.Vuong_Sonah_2015
Markers associated with Soybean Cyst Nematode (SCN) resistance in Soybean, relative to the Wm82.gnm2 assembly. Markers derive from the SoySNP50K set.
Wm82.gnm2.mrk.Zhao_Teng_2017
SLAF-seq markers associated with resistance to soybean cyst nematode (SCN) HG type 2.5.7, with respect to Wm82 gnm2 coordinates (derived via sequence homology, from markers originally with positions in Wm82 gnm1).
Wm82.gnm4.mrk.SoySNP50K
SoySNP50K is an Illumina Infinium II BeadChip that contains over 50,000 SNPs from soybean useful for genotyping a wide variety of cultivated and wild soybean accessions.
Wm82.gnm6.mrk.SoySNP50K
SoySNP50K is an Illumina Infinium II BeadChip that contains over 50,000 SNPs from soybean useful for genotyping a wide variety of cultivated and wild soybean accessions.

Synteny

Wm82.gnm2.syn.HXNY
Synteny mappings among the genomes. See genome assemblies indicated in the synteny comparisons.

Diversity

Wm82.gnm1.div.ContrerasSoto_Mora_2017
Haplotype information for 169 Brazilian soybean cultivars genotyped with the 6K SNPchip.
Wm82.gnm1.div.Hu_Zhang_2020
Variant information for 96 Glycine soja accessions, genotyped using the NJAU 335K SoySNP array, provided by Guizhen Kan.
Wm82.gnm1.div.Song_Hyten_2015
VCF, HapMap, and Flapjack files containing genotype information of the whole U.S. soybean germplasm collection using the SoySNP50K BeadChip.
Wm82.gnm1.div.Song_Wei_2015
The polymorphism information of INDELS and SNPs between Hedou12 and Wm82.
Wm82.gnm1.div.Song_Yan_2017
Haplotypes of SoyNAM parents and progeny with respect to Williams 82 genome assembly gnm1.
Wm82.gnm1.div.Wang_Jiao_2016
Variant information in vcf format for 367 Chinese soybean accessions using the NJAU 355K SoySNP array, provided by Dr. Jiao Wang.
Wm82.gnm1.div.Zhou_Jiang_2015
Resequencing information for 302 wild and cultivated soybean accessions.
Wm82.gnm2.div.Jeong_Moon_2019
Variant information for 4234 soybean accessions (Glycine max and Glycine soja) genotyped with the Axiom SoyaSNP array containing 180,961 SNPs.
Wm82.gnm2.div.Lam_Xu_2010
VCF file containing genotype information for 31 wild and cultivated soybeans received from Meng Ni, Tin Hang, and Hon-Ming Lam.
Wm82.gnm2.div.Lee_Jeong_2015
Text, VCF, HapMap, and Flapjack file containing genotype information for 222 Korean accessions.
Wm82.gnm2.div.Liu_Shi_2025
Predicted SNP effects relative to genes in Williams 82 gnm2 (Wm82.a2.v1).
Wm82.gnm2.div.Song_Hyten_2015
VCF, HapMap, and Flapjack files containing genotype information of the whole U.S. soybean germplasm collection using the SoySNP50K BeadChip.
Wm82.gnm2.div.Song_NAM_2021a
Imputed genotypes for 5176 Nested Association Mapping RILs relative to 40 NAM parental genotypes, genotyped using the SoySNP50K and SoyNAM6K assays. These data sets report alleles at 29,417 SNP positions.
Wm82.gnm2.div.Song_NAM_2021b
Imputed genotypes for 5176 Nested Association Mapping RILs relative to 40 NAM parental genotypes, genotyped with WGS and the SoySNP50K and SoyNAM6K assays. These data sets report alleles at 423,419 SNP positions.
Wm82.gnm2.div.Song_Yan_2017
Haplotypes of SoyNAM parents and progeny with respect to Williams 82 gnm2.
Wm82.gnm2.div.Torkamaneh_Laroche_2017
Resequencing information for 102 Canadian soybean accessions.
Wm82.gnm2.div.Torkamaneh_Laroche_2019
HapMap Data, GmHapMap, for 1007 accession of Glycine max and Glycine soja, from Torkamaneh et al. (2019)
Wm82.gnm2.div.Valliyodan_Brown_2021
Variant information for 481 soybean accessions (Glycine max and Glycine soja), provided by Babu Valliyodan and Henry Nguyen, as part of the United Soybean Board project to genotype and characterize diverse accessions.
Wm82.gnm2.div.Wei_Mesquita_2017
VCF file containing GBS information for 374 U.S. and Brazilian accessions against 68301 SNPs.
Wm82.gnm2.div.Wickland_Battu_2017
VCF files containing genotype information for 3 populations of soybean produced by GB-eaSy.
Wm82.gnm2.div.Zhang_Jiang_2020
Variant information for 1,556 resequenced soybean accessions Glycine max and Glycine soja, provided by Yong-Qiang An and Rick Meyer (Zhang et al., 2022), relative to Wm82 assembly 2.
Wm82.gnm2.div.dosSantos_Valliyodan_2016
VCF and HapMap file containing genotype information for 28 Brazilian soybean accessions.
Wm82.gnm4.div.Song_Hyten_2015
VCF and HapMap files containing genotype information of the whole U.S. soybean germplasm collection using the SoySNP50K BeadChip.

Glycine soja

Genomes

F_IGA1003.gnm1.V9RB
Genome assembly files for Glycine soja F (F_IGA1003 in publication; WHFS_GsojaF_1.0 in the GenBank assembly record)
PI483463.gnm1.YJWS
Glycine soja accession PI 483463 genome assembly, v1.0; JGI name Glycine soja v1.1
PI_549046.gnm1.TSLF
Genome assembly for Glycine soja accession PI_549046 (SoyW02)
PI_562565.gnm1.Z2VG
Genome assembly for Glycine soja accession PI_562565 (SoyW01)
PI_578357.gnm1.1XT1
Genome assembly for Glycine soja accession PI_578357 (SoyW03)
W05.gnm1.SVL1
Genome assembly files for cultivar W05 from Xie, Lam et al. (2019): A reference-grade wild soybean genome

Annotations

F_IGA1003.gnm1.ann1.G61B
Genome annotation files for cultivar F
PI483463.gnm1.ann1.3Q3Q
Glycine soja accession PI483463 genome annotation files; JGI name Glycine soja v1.1
PI_549046.gnm1.ann1.65KD
Gene annotations for Glycine soja accession PI_549046 (SoyW02)
PI_562565.gnm1.ann1.1JD2
Gene annotations for Glycine soja accession PI_562565 (SoyW01)
PI_578357.gnm1.ann1.0ZKP
Gene annotations for Glycine soja accession PI_578357 (SoyW03)
W05.gnm1.ann1.T47J
Glycine soja accession W05 genome annotation files

Glycine cyrtoloba

Genomes

G1267.gnm1.YWW6
Genome assemblies for Glycine cyrtoloba, accession G1267

Annotations

G1267.gnm1.ann1.HRFD
Genome annotation files for Glycine cyrtoloba, accession G1267

Glycine dolichocarpa

Genomes

G1134.gnm1.PP7B
Genome assemblies for Glycine dolichocarpa, accession G1134

Annotations

G1134.gnm1.ann1.4BJM
Genome annotation files for Glycine dolichocarpa, accession G1134

Glycine falcata

Genomes

G1718.gnm1.B1PY
Genome assemblies for Glycine falcata, accession G1718

Annotations

G1718.gnm1.ann1.2KSV
Genome annotation files for Glycine falcata, accession G1718

Glycine stenophita

Genomes

G1974.gnm1.7MZB
Genome assemblies for Glycine stenophita, accession G1974

Annotations

G1974.gnm1.ann1.F257
Genome annotation files for Glycine stenphyta, accession G1974

Glycine syndetika

Genomes

G1300.gnm1.C11H
Genome assemblies for Glycine syndetika, accession G1300

Annotations

G1300.gnm1.ann1.RRK6
Genome annotation files for Glycine syndetika, accession G1300

Glycine D3-tomentella

Genomes

G1403.gnm1.CL6K
Genome assemblies for Glycine D3 tomentella, accession G1403

Annotations

G1403.gnm1.ann1.XNZQ
Genome annotation files for Glycine D3 tomentella, accession G1403